It would be very convenient if the individual neurons of artificial neural networks corresponded to cleanly interpretable features of the input. For example, in an “ideal” ImageNet classifier, each neuron would fire only in the presence of a specific visual feature, such as the color red, a left-facing curve, or a dog snout. Empirically, in models we have studied, some of the neurons do cleanly map to features. But it isn't always the case that features correspond so cleanly to neurons, espec...